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While Kaede and Muramuko rush away with Susana-O, trees all over Japan begin to wither and die, causing a national uproar. Meanwhile, at the resort of Nara, a new type of Aragami are terrorizing the land by polluting the air with toxic pollens. Disheartened Momiji, distracted Yaegashi and the rest of the TAC team set out to destroy the pollen monsters with their newest invention.
